Category Archives: Plant Profile

11 June 2015 –
Eastern Prickly Pear –
Opuntia cespitosa

DSC00712This afternoon, I took a quick jaunt out to Lake Allen to take some pictures of the  Eastern Prickly Pear, a type of cactus that is found on the Refuge. Its scientific name is Opuntia cespitosa Raf. Its natural range is primarily west of the Appalachians and east of the Mississippi. However, there are scattered populations of this species along the Atlantic Seaboard from New England to Georgia.

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The population on the refuge is rather large, covering several acres. It could be one of the largest populations of cactus in Maryland. On the refuge, it thrives in the sandy soils of the northeastern part of the North Tract by Lake Allen and by the firing ranges.

Allegheny Mound Ant mound in the upper left hand corner
Allegheny Mound Ant mound in the upper left hand corner

It is shade intolerant and thrives in full sun, and needs well-drained soil.

DSC00695The fruit of the Opuntia cespitosa is edible, and can stay on the plant until the next spring. Here on the refuge, it blooms from late May to mid-June.

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In a couple of the shots I had to deal with the pesky Allegheny Mound Ants (Formica exsectoides). Without looking, I plopped down on a swarm of them, and within a minute or so they were crawling all over me. I even had to take one off of my forehead. They are an interesting critter and merit a separate blog entry. You can see a couple of their mounds in the pictures on this page. (More about them in future blog entry)

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9 June 2015 – Maleberry – Lyonia ligustrina

(Photo of a maleberry shrub in habit.)

Previous year's seed capsules.
Previous year’s seed capsules.

Maleberry (Lyonia ligustrina (L.) DC.) is an Ericad (member of Ericaceae – Heath Family) that is found on the refuge around the margins of wetlands. Its white flowers mature into brown dry capsules which contain seeds. In the USA, it is found from Maine to Georgia on the East, and Kentucky, Tennessee, and Alabama on the West.

Leaves and seed capsules.
Leaves and seed capsules.

 

 

This specimen was found next to a swampy area under the BGE powerline right-of-way on the North Tract.

Inflorescence
Inflorescence
Inflorescence
Inflorescence

8 June 2015 – Carex amphibola – Eastern Narrowleaf Sedge

(Click on the thumbnails below for larger views of the photos.)
DSC00640 This keys out to Carex amphibola Steud. or Eastern Narrowleaf Sedge. I originally keyed it out to Carex willdenowii, but Wes Knapp suggested that this could be Carex amphibola instead. It was found in a lot that was disturbed in the past. This species had taken over a large area of the lot. The Flora of Virginia states that species is found in moist old fields among other habitats, and this site fits the bill of a moist old field. The location is Hanover Road and Race Road just inside Howard County, Maryland.

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9 May and 2 June 2015 – Which Carex is this?

Specimens of immature plants were collected on 9 May 2015.   I went back to the same site on 2 June and collected some specimens of more mature plants. Maybe they are now mature enough to identify. Any idea?  I believe they may be Carex longii.

(Click on the thumbnails for a full-sized view)

2 June 2015
2 June 2015
2 June 2015
2 June 2015

 

2 June 2015
2 June 2015
2 June 2015
2 June 2015

 

9 May 2015
9 May 2015
9 May 2015
9 May 2015

 

9 May 2015
9 May 2015

1 June 2015 – Swamp Dewberry – Rubus hispidus

DSC00441Today, while on South Tract of the refuge I found another species of Dewberry. It is known as Swamp Dewberry (Rubus hispidus L.) and is also called Bristly Dewberry due to its presence of numerous bristles throughout the plant. The prickles that are present are generally less than 2 to 4 mm long and weak. Swamp Dewberry may be a slight misnomer, although it is frequently found in swampy habitats, it is also found in drier habitats as well. Its native range is Central and Eastern North America from Ontario and the Maritime Provinces on the North to Texas and South Carolina on the South.

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Besides have more bristles than prickles, it differs from the other two dewberry species known to occur on the Refuge – Northern Dewberry (Rubus flagellaris) and Southern Dewberry (Rubus trivalis) with its leaflet tips being blunter/more obtuse and more flowers and later fruits per flowering stem.  Its flowers are smaller on the average than the other dewberries.

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They are in full bloom on the refuge in open areas such as the powerline right-of-ways and in swales and ditches along side roads. It can also be found in wet woods.

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23 May 2015 – Staggerbush – Lyonia mariana

(Photo above: close up of inflorescence.)

Inflorescence
Inflorescence

 

 

This morning I visited the North Tract’s Pitch Pine Barrens area and found a fairly good-sized patch of Staggerbush in full bloom. The showy flowers really stand out. The scientific of this ericad (member of the Heath Family – Ericaceae) is Lyonia mariana (L.) D. Don. Its native range is split in two. On the US East Coast, it is found from Southern New England to Florida, and again from Missouri to Texas. It is not found in between the two ranges.

 

 

Shrub in habit.
Shrub in habit

Staggerbush is tolerant of a wide range of habitats from wet and boggy sites to well drained but wet sandy locations. As it is a constituent of the New Jersey Pine Barrens, it is not surprising to find it on the Pine Barrens on the Refuge. Its fruit is dry and splits open when ripe.

Dried fruits
Dried fruits from previous year
Flowers and fruits
Flowers and fruits

 

 

 

 

 

 

 

 
REFERENCES:

Gleason, Henry and Arthur Cronquist. 1991. Manual of Vascular Plants of Northeastern United States and Adjacent Canada. D. Van Nostrand Company. New York, New York. 910 pp.

Natural Resources Conservation Service, United States Department of Agriculture. Entry for Lyonia mariana. http://plants.usda.gov/core/profile?symbol=LYMA2

16 May 2015 – Cut-leaf Evening Primrose – Oenothera laciniata

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The Cut-leaf Evening Primrose is in its prime on the refuge. It is native to most of North America, except for the far western regions. Its scientific name is Oenothera laciniata Hill.

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In some southern states it is considered invasive. It generally occurs in areas that have been disturbed by man.  You can find it on the refuge in open areas and old fields.  However, you may have to look carefully to find it because it occurs in small numbers hidden amongst the grass and other vegetation.

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It and other evening primroses bloom at night and close up as the sun rises in the sky.  This adds to its inconspicuousness.

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16 May 2015 – Northern Dewberry – Rubus flagellaris

_DSC0642The northern dewberry (also known as the common dewberry) is in full bloom on the refuge right now. Its accepted scientific name is Rubus flagellaris Willd. It is native to eastern North America from the Atlantic Coast on the east to Ontario, Minnesota, and Texas on the west. Botanists call it a liana, or a woody vine that does not support its own weight.

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One of the distinguishing traits for this dewberry are the stout-based, usually recurved prickles on the stems. It is also distinguished from other trailing species of Rubus or dewberries by its few-flowered inflorescences with ascending pedicels (flowering stems) and glabrous to sparsely pubescent leaves.

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It generally prefers drier or mesic habitats and open sunny locations, but can sometimes be found in wet habitats.  It can be seen on the refuge in open ranges and old fields. It is especially abundant in the power line right-of-ways.
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The berries are edible to humans but are hard to find because they are relished by small mammals which get to them first. Deer also browse on the stems.

15 May 2015 – One-flowered Cancer root – Orobanche uniflora

_DSC0628Earlier this week, on the Maryland Biodiversity Project pages I saw a couple of photos of One-flowered Cancer root (Orobanche uniflora L.) on the North Tract taken by Richard Orr. Wow, since I have not seen Cancer root on the North Tract, it had to be documented for the record.  Well, this afternoon, Richard kindly showed me the location along the Patuxent River where he found them. (You can see the photos I took on this page. Click on the thumbnails for a full-sized view.)

One-flowered Cancer root is holoparasitic and lacks chlorophyll; it is dependent on the host plant for nutrients. Its hosts includes members of the Asteraceae and the Saxifragaceae. It is found throughout the USA, except Hawaii, the southern tier of Canadian provinces.

It is also called One-flowered Broomrape, Ghostpipe, and Naked Broomrape. Interestingly, although it is native to the USA, the Federal Government and nine states consider this to be a noxious weed.

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11 May 2015 – Deerberry – Vaccinium stamineum

Close up of flowers
Close up of flowers

This afternoon, I went looking for bogs that I had not found yet.  On the edge of one of the bogs were some Deerberry (Vaccinium stamineum) bushes.  They were putting on a showy display so I decided to photograph them.

 

 

Almost ripe berries (photo taken in July 2011)
Almost ripe berries (photo taken in July 2011)

 

Deerberry is a member of the genus Vaccinium which features blueberries, cranberries, and in the Pacific Northwest some types of huckleberries. It is native to eastern North America from Maine to Texas, and can be found in Ontario, Canada. Its fruits are edible to humans. They are readily eaten by various animals. Its twigs and foliage is relished by deer.

Once established, a single plant can form a large clonal thicket.  I just think it is a beautiful plant when in bloom.

 

Upper side of leaf
Upper side of leaf
Lower side of leaf
Lower side of leaf

 

Cluster of flowers
Cluster of flowers

25 April 2015 – Primrose-leaved Violet – Viola primulifolia L.

(Photo above: a picture of a plant with the rhizomes clearly visible.)

A look at the inner part of the flower.  You can see the beard hairs on the lateral petals and the purple lines on the lower petal.
A look at the inner part of the flower. You can see the beard hairs on the lateral petals and the purple lines on the lower petal.

 

Today I had the privilege of taking Bill Hubick and Michael Ostrowski on an abbreviated tour of the North. Along the way, we found this cute little plant called a  Primrose-leaved Violet (Viola primulifolia L. (pro sp.) [lanceolata × macloskeyi]) today along the southern edge of Burr Reed Swamp.

 

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This small-flowered species of violet is found in eastern North America along the east coast from Maine to Florida and to Minnesota to Texas on the west. It likes to have its feet wet, being found in swampy and boggy sites. Its flowering stem is borne on a leafless stalk that rises from the root crown.

 

The hairless orbicular shaped leaf.
The hairless orbicular shaped leaf.

 

 

The leaves are glabrous, which to a botanists means the surface is without hairs or projections. They are orbicular to orbicular-ovate in shape, which means the outline of the leaves resemble an orbit.

 

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Plants spread vegetatively via creeping rhizomes with stolons (prostrate stems) and are capable of producing sizable colonies.

 

Full view of the flower.
Full view of the flower.

The flowers are only 1/4 to 3/8 inch across. The lateral petals are bearded and lower petal has purple lines.

 

 

 

 

 

Below are some random shots of other Viola primuifolia plants found on the random. Click on each for the full sized view.

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23 April 2015 – Blue Marsh Violet

DSC00082 close upToday, I drove down Telegraph Road stopped by a swamp about a 1/2 mile south of Wildlife Loop. The swamp is called a Southern New England/Northern Piedmont Red Maple Seepage Swamp (USNVC: CEGL006406). In the swamp, I found a bunch of violets which turned out to be a Blue Marsh Violet (Viola cucullata Aiton).

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This violet’s most distinctive characteristic is the club-shaped (clavate) hairs on the lateral petals, which you can see in the photo here. It is native in North America from Minnesota to Ontario to Newfoundland on the north to Georgia in the South. It likes wet boggy soil just like where I found it on the refuge.

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Violas can be divided into two different types, acaulescent and caulescent. In acaulescent species the flowers are born on a stem that rises directly from the root crown. In caulescent species, the flowers are born on a leafy stem.  The Blue March Violet is an acaulescent species.

 

CITATIONS:

Brown, M.L. and R.G. Brown. 1984. Herbaceous Plants of Maryland. Port City Press, Baltimore, MD.

Gleason, H. A. and A. Cronquist. 1991. Manual of the Vascular Plants of Northeastern United States and Adjacent Canada. 2nd ed. New York Botanical Garden, NY.

McKinney, L.E. 1992. A taxonomic revision of the acaulescent blue violets (Viola) of North Am. Sida Botanical Miscellany, 7: 1–59

Nature Serve. Ecological Association Comprehensive Report – http://explorer.natureserve.org/servlet/NatureServe?searchCommunityUid=ELEMENT_GLOBAL.2.688675 Accessed 23 April 2015.

Nature Serve. Viola Cucullata Comprehensive Report. http://explorer.natureserve.org/servlet/NatureServe?searchName=Viola+cucullata Accessed 23 April 2015.

Weakley, A.S. 2014. Flora of the southern and mid-Atlantic states. University of North Carolina Herbarium, North Carolina Botanical Garden, University of North Carolina, Chapel Hill.

17 April 2015 – Cinnamon Fern Fiddleheads

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Today while slogging around in the magnolia bog I came upon some Cinnamon Fern (Osmundastrum cinnamomeum (L.) C. Presl.) fiddleheads. They look ghostly to me because they are mostly white with a little bit of green and brown mixed in. Many people believe that fiddleheads are edible. The fact is that only a few species are really good to eat. In the case of cinnamon fern, their fiddleheads are mildly toxic. Some people probably could eat a small amount without any ill effects, but the more you eat, the higher the possibility of getting ill.

The Cinnamon Fern has worldwide distribution. It is found in far flung locations such as Korea, Japan, China, southeastern Siberia, Thailand, Myanmar, and Vietnam. It occurs in South America south to Peru and Paraguay. In North America, it is found in the Eastern United States, in Canada from Ontario to Labrador, and in Mexico and the West Indies.

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You can bet I will be taking some pictures of this lovely plant later in the season.

16 March 2014 – Quirky Hybrid Oaks of Patuxent

Did you know that the Patuxent Research Refuge is the home to several types of hybrid oaks? Read on to learn more.

20140316bThe oak-rich Patuxent Research Refuge has 16 native oak species, one naturalized exotic oak species, and at least ten types of trees that are regarded as oak hybrids. The reason so many oaks species call the Refuge their home is due to the Refuge location near the Fall Line between the Coastal Plain and the Piedmont and at the same time, it is at a crossroads between northern and southern species of oak.

In North America, there are two basic groups or sections of oak species, the red oaks and white oaks. An easy way to tell them apart is determine whether or not there are bristles at end of the lobes of the leaves. Red oaks have bristles while white oaks do not. When considering hybrid oaks, it is important to distinguish between these two groups of oaks because red oak species will cross only with other red oak species and white oak species will cross only with other white oak species.

Hybrid oaks may be found anywhere two parent species are found. However, the Refuge seems to have more than its share of hybrids when compared to other areas, with at least ten types of hybrids identified so far on the Refuge. Undoubtedly, there are other types of hybrids on the Refuge which have not yet been identified. The reason for the richness of hybrid oaks on the Refuge is probably due to the influence of human activity.

Under normal conditions, there are barriers against hybridization. These include pollination timing, closeness to other trees of the same species, and the rejection of “foreign” pollen by the female (pistillate) flowers. While hybridization occurs naturally, many of these barriers can be easily broken down and the process accelerated because of human disturbance. This phenomenon is easily observed on the Refuge due to the Refuge’s long history of farming, logging, and more recently, military training. These activities have contributed to an increased number of opportunities for individual trees to become isolated and marginalized from others of their own species. When this happens, these isolated trees are overwhelmed with pollen from nearby trees of a closely related species – under normal circumstances they would not readily cross. It is interesting to note that this means the mother tree of the hybrid can generally be assumed to be the tree that was isolated from others of its species.

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There are several young Fernow’s Oak (Quercus ×fernowii) trees at one location along Wild Turkey Way. (The Fernow’s Oak is the hybrid between the Post Oak (Quercus stellata) and the Common White Oak (Quercus alba.)) Here, a mature Post Oak which found itself isolated from other Post Oaks but surrounded by a large number of Common White Oaks. As a result, the pollen from the surrounding Common White Oaks fertilized the female flowers of the mother Post Oak and the progeny of this tree are found nearby. As with most other hybrid oaks, mature Fernow’s Oak can produce viable acorns. However, none have been noted here yet.

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Another easily observed and notable hybrid specimen is a Saul’s Oak tree located along Wild Turkey Way near Snowden Cemetery. The Saul’s Oak (Quercus ×saulii, a cross between the Common White Oak and the Chestnut Oak (Quercus prinus.)) This particular tree is a known prolific acorn producer. The mother tree is probably a Chestnut Oak based on the fact that there are a large number of Common White Oaks at this location.

You can see a short article on Saul’s Oak on the refuge by clicking here.

Sometimes it is difficult, if not almost impossible, to determine the parentage of some individual trees which appear to be hybrids. These trees may clearly exhibit some, but not all, of the characteristics of one parent species, but not the other. There are several trees on the Refuge that fall into this category. It might be possible to clear this up by DNA testing.

Then there is the process called introgression or hybrid swarming. This happens when a viable hybrid will backcross with one or another of the parent species creating progeny that are not clearly intermediate between the two parents. The observer is forced to speculate on the parentage of the tree based on what species of trees are located nearby. If backcrossing with the parents, other hybrids, and backcrossed trees continues, trees exhibiting a highly variable range of characteristics between the two parents can result. There is evidence that this is also happening on the Refuge.

Finally, it is possible for hybrids to cross with a third species or even hybrids from different sets of parents to cross with each other. The resulting individual trees have three or more “parents.” This is not yet known to happen on the Refuge, but has been well documented as happening in other locations like California.

If there is interest, the Refuge may schedule a walk focusing on oak hybrids in the future.

PATUXENT RESEARCH REFUGE OAKS –

Hybrids –
Bush’s Oak – Quercus ×bushii [marilandica × velutina] –
Fernouw’s Oak- Quercus ×fernowii [alba × stellata] –
Bartram’s Oak – Quercus ×heterophylla [phellos × rubra] –
A hybrid oak – Quercus xhumidicola
A hybrid oak – Quercus ×incomita [falcata × marilandica] –
Jack’s oak – Quercus xjackiana
Saul’s Oak – Quercus ×saulii [alba × montana] –
Subfalcate Oak – Quercus ×subfalcata [falcata × phellos] –
Willdenow’s Oak – Quercus ×willdenowiana [falcata × velutina]

Native Red Oaks –
Scarlet Oak – Quercus coccinea
Southern Red Oak – Quercus falcata
Bear Oak – Quercus ilicifolia
Shingle Oak – Quercus imbricaria
Blackjack Oak – Quercus marilandica
Pin Oak – Quercus palustris
Willow Oak – Quercus phellos
Northern Red Oak – Quercus rubra
Black Oak – Quercus velutina

Native White Oaks –
Common White Oak – Quercus alba
Swamp White Oak – Quercus bicolor
Overcup Oak – Quercus lyrata
Swamp Chestnut Oak – Quercus michauxii
Chestnut Oak – Quercus montana
Dwarf Chinkipin Oak – Quercus prinoides
Post Oak – Quercus stellata

Naturalized Exotic Oak –
Sawtooth Oak – Quercus acutissima

Planted Not Established Exotic Oak –
Bur Oak – Quercus macrocarpa

5 April 2013 – Carey’s Balsamroot – Balsamorhiza careyana

Carey's Balsamroot - full flower head
Carey’s Balsamroot – full flower head

Carey’s Balsamroot (Balsamorhiza careyana A. Gray) is found on east of the Cascade Mountains in Washington and Oregon. This member of the Aster Family (Asteraceae) is not as widespread as the Arrowleaf Balsamroot, but is common in its more limited range. The leaves are similar in shape to the Arrowleaf Balsamroot, but not as hairy and the margins are more wavy. The plants pictured here were growing at the Vantage Overlook next to Interstate 90 along the Columbia River Gorge.

It thrives in a wide variety of habitats, but seems to like drier locations than its relative Arrowleaf Balsamroot. The plant was named for Asa Gray’s botanist friend John Carey.

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Carey’s Balsamroot –
full plant
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Carey’s Balsamroot
Carey's Balsamroot - Top side of leave
Carey’s Balsamroot – top side of leaf
Underside of leaf Carey's Balsamroot
Underside of leaf
Carey’s Balsamroot

5 April 2013 – Big Sagebrush – Artemisia tridentata

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Plants in habit

Big Sagebrush (Artemisia tridentata Nutt.) is an aromatic evergreen shrub from the Aster Family (Asteraceae). It is native to Intermountain West of North America, thriving in arid and semi-arid habitats ranging from cold deserts to dry mountain slopes. It is the dominant species in large swaths of its natural range.

It is a coarse shrub that combines a taproot and a shallow root system so that it can take advantage of both watertable water and any available surface water. Individual plants can live to be well over 100 years.

Leaf showing the characteristic three terminal lobes

It has pale-yellow flowers and silvery-grey foliage. The leaves are between 3/8 to 1-1/4 inches long and have a three lobes on their tips.

The plant contains numerous aromatic and volatile oils (camphor, terpenoid oils, etc.) which gives it its characteristic odor and bitter taste. The bitter taste serves to discourage browsing by herbivores.

Big Sagebrush

Habitat alteration is reducing its range. Big Sagebrush is not fire tolerant and destruction of native grasses and herbs predominantly by overgrazing creates conditions for fire which in turn creates an opening for invasives such as Cheatgrass to takeover. This makes it difficult for Big Sagebrush to re-establish itself.

5 April 2013 – Western Tansymustard – Descurainia pinnata

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Plants in habit

Descurainia pinnata (Walter) Britton is a member of the Mustard Family (Brassicaceae) known as the Western Tansymustard. It is an annual native to North America and is found in every USA state except Alaska, Hawaii, and Alabama as well as throughout Canada and Mexico. It can be found in a variety of habitats and can springs up in barren sites with poor soil.

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Inflorescence from above

It especially seems to do the best in the drier plant communities like sagebrush deserts and short bunchgrass communities. This species is highly variable with several subspecies, and as far as I can determine, these plants appear to be Descurainia pinnata (Walter) Britton ssp. nelsonii (Rydb.) Detling or Nelson’s Tansymustard .

 

Pinnate leaf

 

The leaves are highly lobed.  At the end of the stems are tiny golden-yellow flowers. The fruits are siliques about 0.25 to 0.75 inches long. The plant is nutritious to grazing animals in small quantities, but can be toxic in large quantities due to nitrates and other related compounds. The seeds are similar to black mustard and can be consumed as food for humans.

These photos were taken at the Vantage Overlook off of Interstate-90.

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Close-up of inflorescence
Side view of inflorescence with siliques starting to form

5 April 2013 – Low Pussytoes – Antennaria dimorpha

Typical mat growing habit of the low pussytoes.

The Low Pussytoes (Antennaria dimorpha (Nutt.) Torr. & A. Gray) is a mat-forming low growing perennial  member of the Aster family (Asteraceae) which prefers drier habitats like sagebrush desert and bunchgrass communities.

It is native to western North America from Southern British Columbia to Northern California to Nebraska. In Washington, it is found in all but two counties east of  the Cascades Crest.

Close up of mature pistillate (female) flowering heads

 

The Low Pussytoes is dioecious, meaning that male (staminate) flowers and female (pistillate) flowers are born on separate plants, or in other words sexually dimorphic. (Thus the species name dimorpha.)

The mats are formed at the top of a many-branched caudex (type of tap root.)

The plant pictured here mature female flowers. Pictures taken at the Vantage Overlook along Interstate 90.

Several flower heads
Granddaughter Nellie pointing out a clump of Low Pussytoes to Grandpa.